Capsids of these viruses are icosahedral, 25 to 30 nm in diam- eter, and are composed of 60 coat protein (cp) subunits [ 10 , 11 ]. the secovirus genome can be monopartite or bipartite. bipartite genomes are divided between two rna segments (rna1 and 2), each modified by the addition of a 3’-termi- nal poly(a) tail and a covalently bound genome-linked viral protein (vpg) at the 5’end. rna1 ranges in length from 6 to 8 kb and encodes the proteins necessary for cytoplasmatic replication. in contrast, rna2 is smaller, ranging from 2 to 4 kb, and encodes the movement protein (mp) and up to three cps [ 8 ]. the pro-pol sequence on rna1 is used to infer phylogenetic relationships within the family and to other members of the order [ ]. currently, members of the family secoviridae are classified into nine genera: como- virus fabavirus nepovirus cheravirus sadwavirus tor- radovirus sequivirus stralarivirus , and waikavirus [ ]. in addition, the genus is divided into three sub- genera, stramovirus satsumavirus cholivirus ]. pineapple secovirus a (psv-a), a recently assigned